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Science Introduction  |  Background  |  Step One  |  Step Two  |  Step Three  |  Next Steps  |  References  |  Maps  



 Making Connections for Wildlife
Aligning Transportation Projects with State Wildlife Action Plans:  A Step-by-Step Guide for Integrated Conservation Planning

Step Two: Identify and Map Wildlife Linkages

The development of a spatially explicit wildlife linkages data layer involves multiple steps, each of which is outlined below. Once complete, these data can be overlaid with spatial data on upcoming transportation projects in the STIP to highlight potential conflicts with wildlife.

Researchers at Northern Arizona University recently developed Corridor Design, a freeware geographic information system (GIS) tool to assist with the linkage design process. This tool was developed to aid in the design of landscape-scale corridors in a heterogeneous environment. The ArcGIS tool encompasses a series of spatial analyses that walk the user through defining suitable habitat for target species, identifying core areas, finally, mapping the optimal corridors between core areas.

While other tools and methodologies are also available and may be adapted to this process, we feel that the Corridor Design tool is the most appropriate for fine-scale mapping of wildlife linkages. Should your team elect to use a different analysis and mapping tool, you may still want to review the steps described in this section, as the general processes and considerations will remain similar regardless of the specific GIS methodology used.

Link: 

Corridor Design Tool Link

GIS Mapping Steps

The following steps should be pursued in close collaboration and communication with the Partnership Team, so that all members of the team – biologists and non-biologists alike – fully understand and participate in the analysis process. These steps are described below, however, for more information, we highly recommend that you also consult the Corridor Design webpage (see link, above).

GIS Mapping Step 1 – Identify Analysis Area(s)

Define what you are trying to connect. Identify and prioritize core and linkage areas. Several states, including Colorado, have completed statewide connectivity assessments, which can serve as the basis for identifying analysis areas for further definition using the Corridor Design tool.

Links:

Report: Inventory of States with Completed or In-Progress Connectivity Assessments

GIS Mapping Step 2 – Compile List of Target Species:

Consider the following types of species:

  • Habitat specialists;
  • Species with limited movement ability, or species that are sensitive to roadways or other types of barriers;
  • Area-sensitive species that require large or well-connected landscapes to maintain a viable population and genetic diversity;
  • Keystone predators, seed dispersers, etc;
  • Species listed as threatened or endangered under the Endangered Species Act;
  • State species of concern.

In reviewing the selected suite of target species, make sure that the range of habitat types and ecological systems present within the analysis area are captured by the target species.

GIS Mapping Step 3 – Define Habitat Parameters and Compile Data Layers

A number of parameters must be defined for each target species to conduct the corridor modeling. Parameter definition should be conducted in close collaboration with each species biologist. Review of the parameters by several biologists is recommended, whenever feasible, and keep in mind that the parameters may require tweaking through several iterations in order to develop appropriate models.

There are four primary factors that are evaluated for each target species: land cover, elevation, topography, and distance from roads. Each of these criteria are weighted (0-100%), depending on the degree to which they influence a given target species’ habitat use.  Each factor is given a percentage of its importance to the species so that the factors add up to 100%.  The factors are then combined spatially to create the final HSM where every pixel has a value of the habitat importance to the species. Then, for each factor with a weight greater than zero the weighted geometric mean was calculated by raising each factor by its weight, and multiplying the factors.

If a particular target species’ habitat use cannot be adequately captured by these four factors, then the team may want to consider adding a new factor for that species (e.g., aspect, soil type). However, the makers of the Corridor Design tool argue, and we concur, that the use of few factors and few categorical metrics within each factor is better than many factors or numerous categories within a factor – complex models become increasingly abstract and are likely to be poor representations of what is actually occurring on the ground.  

Other parameters that must be defined for each target species include minimum core patch size and daily dispersal distance. In addition, the team must determine how start and end points for each linkage analysis will be determined. In Arizona, protected area were used as start and end points, whereas in Colorado, in most cases, suitable habitat was used to define start and end points.

The primary data layers needed for the Corridor Design Analysis include: land cover (reclassified to a smaller number of habitat groupings), digital elevation model (30m resolution), and roads. Additional data layers may be needed if other factors were identified as primary influences on a given target species’ habitat preferences.

GIS Mapping Step 4 – Adapt Corridor Design Tool

The first step of adapting the Corridor Design tool is to define the areas of focus. More specifically, to identify areas where species are moving or potentially are moving through the landscape. Once the analysis area has been selected then the habitat suitability model can be clipped to the specific area for development of the habitat suitability model.  The next step is to identify potential breeding patches and potential population cores. 

Potential breeding patches = areas large enough to support one breeding event for one season. 

Potential population cores = areas large enough to support a breeding population for approximately ten years.

To create the Habitat Suitability Model (HSM) for each species, each analysis cell of the land cover data layer is weighted based on the habitat preferences as defined by the species experts (Fig. 2).

Land Cover Weights:

1-3 = strongly preferred (1 is best)

4-5 = usable but suboptimal habitat

-------------------------------------------------------------  Suitability Threshold

6-7 = not breeding habitat, but occasionally used

8-10 = strongly avoided (10 is worst) 

The suitability of the habitat is evaluated such that areas above the habitat threshold (i.e., weighted < 5) are joined into polygons of suitable habitat. Core habitat patches are created by combining all suitable and unsuitable habitat that falls within the designated daily dispersal distance, such that unsuitable habitat within a matrix of suitable habitat may be included in a core habitat patch. Suitable habitat beyond the species’ dispersal distance is not included in a core patch. Each habitat patch is then evaluated based on its size, relative to the minimum patch size needed to support a breeding event and the minimum patch size needed to support a core population for at least ten years – habitat patches smaller than these minimums are not considered core habitat.

Figure 2: Example of a Habitat Suitability Model.

Protected areas, species data on existing breeding populations or occupied habitat, or potential core habitat patches (derived from the HSM) are used to define the endpoints (actually areas) for the linkage analysis model. These endpoints define the area between which the linkage will be identified.

The Corridor Design analysis is performed creating a cost surface based on the HSM (Fig. 3). This is similar to a least-cost path analysis, but also incorporates suitable habitat patch sizes (i.e., small, stepping-stone patches that are too small to be core habitat patches, but may still be important as animals move between core habitat patches) and the species’ daily dispersal distance threshold. The resulting movement surface is a gradation that demonstrates the difficulty (i.e., cost) of moving between the two identified endpoints.

Figure 3: Example of a cost-surface.

Based on this gradated movement surface, the next step is to determine an appropriate linkage width – this width is essentially a cut-off that defines the linkage area. The Corridor Design tool recommends creating slices of the lowest cost 0.1 to 10 percent of the total analysis area to define the actual linkage area (Fig. 4). This width requirement should be reviewed by the species experts.

The Corridor Design tool also provides the ability to combine individual species linkages into a multi-species linkages. In many cases, individual species linkages may overlap, highlighting the importance of specific portions of the linkage area for more then one modeled species. A multispecies approach is also recommended for indicating where appropriately designed mitigation can benefit multiple species.

GIS Mapping Step 5 – Biological review of species linkage models

A final review of the linkage models by the species experts is necessary to ensure that the linkages properly represent the species movement needs as accurately as possible and to determine whether any adjustments need to be made to the model inputs. 

Figure 4: Example of linkages between two defined core habitat patches, with the recommended range of 0.1-10% of the analysis area displayed to define linkage width.

Colorado Case Study

GIS Mapping Step 1 – Identify Analysis Area(s)

The Colorado Team had the benefit of a statewide connectivity assessment to serve as the basis for the Corridor Design analysis. Linking Colorado’s Landscapes, completed in 2005, was a comprehensive effort to identify broad-scale movement corridors for Colorado wildlife through a process involving a series of expert workshops and computer modeling (Fig. 5). The project was conducted as a partnership between the Southern Rockies Ecosystem Project, the Colorado Department of Transportation, the Federal Highway Administration, the Nature Conservancy, and Colorado State University. This assessment resulted in the identification of 176 wildlife linkages across the state. These linkages – represented as coarse movement arrows – provided the analysis areas for linkage definition in the Corridor Design process.

GIS Mapping Step 2 – Compile List of Target Species:

The Partnership Team identified a diverse set of target species to capture the range of ecological and topographic conditions present in the state, with the intention of

Figure 5: High priority linkages identified in Colorado’s statewide connectivity assessment. A total of 176 linkages were identified and prioritized across the state.

creating a linkage design that would ensure their long-term viability. Target species met one or more of the following conditions: species identified as Tier 1 or Tier 2 species in the Colorado State Wildlife Action Plan (see box), species that are frequently involved in animal-vehicle collisions, and/or species of special management concern to CDOW (e.g., at-risk species, game species).

Target Species

Justification

Bighorn sheep

CSWAP Tier 2 species

Black bear

Species of management concern

Black-tailed prairie dog

CSWAP Tier 1 species

Boreal toad

CSWAP Tier 1 species

Canada lynx

CSWAP Tier 1 species

Elk

Species of management concern; AVC concern

Gray wolf

CSWAP Tier 1 species

Gunnison’s prairie dog

CSWAP Tier 1 species

Meadow jumping mouse

CSWAP Tier 1 species

Mountain lion

Species of management concern

Mule deer

Species of management concern; AVC concern

Pronghorn

Species of management concern; AVC concern

Swift fox

CSWAP Tier 1 species

White-tailed prairie dog

CSWAP Tier 1 species

Notably, there are several species that were identified as target species at the outset of the project, but were later removed from the list of target species being modeled. For example, meadow jumping mouse is a riparian species restricted to the Front Range of Colorado. Given its habitat is naturally linear and restriction to the riparian zone, linkages for this species cannot be mapped in the same manner as for other species. Instead, it is appropriate to highlight all suitable habitat for this species as a ‘conflict zone’ where special consideration for avoidance and mitigation measures is needed to adequately protect the species’ ability to move through its habitat. Similarly, species with small-scale movements, such as herpetofauna and some small mammals might operate at too fine of a scale to have their movement patterns adequately captured using this modeling process. In these cases, suitable habitat may be a better determinant of where avoidance or mitigation measures are needed. Attempts were made to model gray wolf and mountain lion, however, it was determined that the parameters for these species were too general to discern linkages between habitat core areas. To adequately model linkages for these species, other factors, such as prey availability, may be better suited for representing these species use of the landscape. Following is a list of species that were determined to be inappropriate for inclusion in the linkage design process:

Target Species

Reason for Not Using Model Analysis to Identify Linkages

Alternatives

Black-tailed prairie dog

Movements too fine-scale to model adequately

Substitute all existing/potential habitat

Boreal toad

Movements too fine-scale to model adequately

Substitute all suitable habitat

Gray wolf

Habitat generalist – models poorly

Consider using prey availability as a surrogate

Gunnison’s prairie dog

Movements too fine-scale to model adequately

Substitute all existing/potential habitat

Meadow jumping mouse

Movements too fine-scale to model adequately

Front Range riparian obligate – substitute all existing/potential habitat

Mountain lion

Habitat generalist – models poorly

Consider using prey availability as a surrogate

White-tailed prairie dog

Movements too fine-scale to model adequately

Substitute all existing/potential habitat

GIS Mapping Step 3 – Define Habitat Parameters and Compile Data Layers

We used the following data layers to define each factor in the habitat suitability models:

Data Layer

Data Source

Land Cover

Southwest Regional GAP Analysis – 102 land cover categories.

Elevation

30 meter resolution National Elevation Dataset (NED)

Topographic Position

Created from the 30-m NED using the Create Topographical Position tool in Corridor Design

Distance to Roads

Buffered distance (by species) from highways, major roads, local roads, and Forest Service roads

List of CDOW biologists Engaged in Model Parameterization:

      

Name

Species Expertise

Jerry Apker

Black Bear, Canada Lynx, Mountain Lion

Janet George

Bighorn Sheep

Tina Jackson

Boreal Toad

Aaron Linstrom

Pronghorn

Eric O’Dell

Black-Tailed Prairie Dog

Pam Schnurr

Gunnison’s Prairie Dog

Scott Wait

Bighorn Sheep, Black Bear, Canada Lynx, Mountain Lion

Bruce Watkins

Bighorn Sheep, Elk, Mule Deer, Pronghorn

 

We found working with the species experts to define parameters for each of the target species to be one of the most time-consuming steps in the process, require significant review of the literature as well as one-on-one collaboration. We found that a number of biologists, being unfamiliar with GIS modeling concepts, were hesitant to provide definitive weights for the factors and factor categories as required to construct the models – there can be significant natural variability in activity and habitat use within a species, and in many cases these details are not well understood. Working with the individual biologists was an educational and iterative process for the Project Team, but was well worth the effort to ensure that the resulting species models accurately reflect habitat use.

Example:

Parameters defined for Colorado’s Target Species

GIS Mapping Step 4 – Adapt Corridor Design Tool

The corridors modeled in this case study were derived from a previous study conducted called Linking Colorado’s Landscapes which gathered together species experts from across the state and identify the highest priority linkages in the state.  The results were sketches and lines of important wildlife linkages, but the study did not define the spatial extents of these linkage areas, thus the spatial mapping of these linkage areas was an important next step for integrating wildlife habitat and movement areas into the transportation planning process.

Habitat Suitability Models (HSM) were created for bighorn sheep, black bear, Canada lynx, elk, mule deer, pronghorn, and swift fox using the process described by the Corridor Design methodology. Potential breeding patches and core habitat patches were then derived from the HSMs. In the case of bighorn sheep, existing populations are well known and do not occur in all areas of suitable habitat.  In this case, data from the Colorado Division of Wildlife on these populations was used to define endpoints for the linkage modeling process.

To best suit Colorado’s needs, the Partnership Team identified several adjustments to the standard Corridor Design methodology: 

  • The objective of the Corridor Design methodology is to define a minimum or ‘biologically best corridor’ between protected lands in order to protect species movement in the face of immediate or impending development. In Colorado, the Project Team decided instead to depict a broader linkage area as a gradation representing the range of ‘costs’ for an animal moving across the landscape between two core habitat patches. Thus, instead of a narrowly defined corridor, the Colorado linkages encompass the concept that some portions of a broader linkage area are more suitable (low cost, or resistance) than others (higher cost, or resistance). This meant that linkage width was not as strictly limited as in the Corridor Design methodology, and that all portions of the linkage do not have equal value to target species.
  • Multi-species linkages were not unioned together as recommended by the Corridor Design process. Instead, the Colorado Team determined that planners can best address the individual and combined needs of a variety of species by having the option of reviewing species linkages individually as well as in an overlay. This allows planners to assess the individual components of the linkage area on a species-by-species basis instead of assuming that all portions of the linkage are equally important for all species considered in the analysis area (Fig. 6a-c).

Figure 6a: Elk linkage at Dallas Divide in southwestern Colorado crossing over Highway 62. Dark brown represents areas of low resistance (low cost of movement) between the two core habitat areas.


Figure 6b: Lynx linkage crossing the same stretch of Highway 62. Note how the low cost (low resistance) portions of the linkage are significantly more restricted for lynx than they are for elk.

Figure 6c: When unioned together, the details of the individual species linkages may be lost. We therefore recommend reviewing the individual species linkages  both separately and combined for a given location for a more complete understanding of where mitigation efforts should be placed for the greatest benefit to multiple species.  

GIS Mapping Step 5 – Biological review of species linkage models

For each target species, we conducted individual meetings with the species experts in which we thoroughly discussed the modeling process, explained each of the inputs into the model, and reviewed the resulting linkage maps. The purpose of these meetings was to evaluate the models and determine if any adjustments were needed to better represent species’ use of the landscape. Each reviewer then responded to the following questions:

  • Based on these results, are the input parameters appropriate for this species (including start/end points, factor weights and factor categorizations?)
  • Are there other factors that should be considered for this species? Such as prey availability, water sources, etc?
  • In the interest of keeping the model simple and straightforward, we did not categorize roads based on traffic volume. How does traffic volume influence movement for this species – should we differentiate among roads types with different traffic volume?
  • Look at how core areas are identified in the Corridor Design process and overlay this information with data on existing wildlife populations. Do core areas defined by the modeling process accurately reflect current population areas?

Questions to Ask

·    Does the State Wildlife Action Plan include data or maps for identifying conservation priorities and important habitat for species of conservation concern?

·    Be clear about the purpose of applying the Corridor Design analysis – what are you trying to connect? Why? Is this the appropriate tool for the job?

·    Does the suite of identified target species sufficiently represent the movement needs of native species in the analysis area?

Key to Success

·    Understand that many biologists are unfamiliar with GIS modeling, and may be hesitant to accept the modeling outcomes. Take the time to explain what the models represent and what they can and cannot do so that the species experts are better equipped to assist in defining habitat parameters.

·    Recognize that the one size may not fit all and that some species are not suited to a GIS modeling process.

·    Actively engage Division of Wildlife GIS staff in the modeling process.

·    Test modeling process through close collaboration with the species experts to fine tune the process and adjust parameters as needed.

·    Develop protocols to ensure that the linkage data is kept up to date. In general, such revisions are unlikely to be needed very frequently, as major changes and new insights into our understanding of a species habitat or habitat preferences are infrequent.



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